（1Institute of Biotechnology and Germplasm Resources，Yunnan Academy of Agricultural Sciences，Key Lab of Agricultural Biotechnology of Yunnan Province，Key Lab of Southwestern Crop Gene Resources and Germplasm Innovation，Ministry of Agriculture，Kunming 650233，Yunnan，China；2Life Science College，Southwest Forestry University，Kunming 650224，Yunnan，China；3Tranditional Chinese Medicine College，Yunnan University of Tranditional Chinese Medicine，Kunming 650500，Yunnan，China）
